2000-2006年 兰州大学 硕博连读自然地理学，理学博士（导师 陈发虎 院士） 
2004-2005年 伦敦大学 Royal Holloway, University College London 第四纪科学硕士 MSc Quaternary Science
1996-2000年 兰州大学 地理学基地班，理学学士

过去环境变化与人类活动、孢粉学与古生态

2018.07-                兰州大学教授、博士生导师
2008.09-2018.06：兰州大学副教授、硕士生导师
2013.07-2014.07：美国伊利诺伊大学和伊州博物馆访问学者 合作导师Eric C. Grimm教授  
2012.12：              挪威卑尔根大学生物系 John Birks教授工作室访问学习 
2008.12-2009.12：法国法国科研中心(CNRS) 近东考古研究所博士后
2007.02-2008.05：兰州大学资源环境学院讲师.

本科生课程
《自然地理学》专业基础课 主讲（2007—） 
《中国与世界地理》全校通识课 主讲（2016—） 
《全球环境变化与人类文明》全校通识课，合讲（2018—）

部分研究生课程
《地理学前沿进展》

	《地球科学进展》期刊专辑客座编辑（2024-2025）
	Environmental Research 期刊专辑客座编辑（2023-2024）
	Frontiers in Earth Science 期刊评审编辑（2022-）
	中国地理学会“人类世与湖泊生态研究工作组”首届委员（2021-）
	中国地质学会“人类世研究分会”首届委员（2022-）
	第四纪科学研究会终身会员，生态系统演变专业委员会委员（2018-）
	中国古生物学会孢粉学会会员
	生态环境部项目评估专家
	教育部人才项目评审专家
	国际第四纪研究会（INQUA）古气候委员会委员
	西部环境教育部重点实验室学术秘书（2015-2018）
	国家自然科学基金、波兰自然科学基金委评审专家
	教育部、中科院学位论文评审专家

针对区域孢粉学现代过程、湖泊岩芯记录、考古遗址地层开展了系统研究，用多种生物与非生物指标重建了北方及西北地区植被、气候等变化历史以及早期牧业活动历史，结合区域人类活动的变化，揭示了气候变化对人群迁徙与文明交流的影响，探究全新世期间人类活动改变生态环境的时限、过程和机理等。围绕上述研究，共发表论文100余篇，论文总引用5200余次（Google Scholar数据）。近5年来，主编著作1部、发表文章60篇（含52篇SCI）；以第一或通讯作者发表文论文36篇（32篇SCI），其中含地学一区文章14篇发表在Nature Communications、Earth-Science Reviews、QSR、GRL、Catena等期刊上。

1. 2014年：教育部自然科学二等奖，亚洲中部干旱区现代间冰期气候和环境变化的“西风模态”，2014.1，教育部，排名第五 
2. 2016年：西部环境教育部重点实验室，管理贡献奖
3. 2018年：兰州大学优秀共产党员
4. 2021年：兰州大学优秀硕士论文指导教师
5. 2022年：兰州大学优秀硕士论文指导教师
6. 2022年：兰州大学优秀博士论文指导教师

在研项目：
1.	国家自然科学基金重大项目课题1: 人类世的历史背景（课题编号41991251），2020.1-2024.12，课题负责人.

结题项目
2.	中科院先导专项A联合攻关项目：高原人-环境相互作用的生态环境影响与调控对策，子课题负责人(子课题编号：XD2009000003) （2018-2022）主持人 280万
3.	国家自然科学基金，重大项目：中国北方干旱半干旱区气候变化及敏感生态系统的响应与适应，课题：中国北方干旱半干旱区森林植被对气候变化的响应(课题编号：41790422)，2018.1-2022.12，参加 
4.	国家重点研发计划：全新世亚洲季风变异与干旱演变及其驱动机制（项目-课题编号：2017YFA0603403）子专题负责人（2017-2022）个人经费80万
5.	科技部重点研发项目，青藏高原科学考察任务六专题一：人类活动历史及其影响（2019-2022）参加，个人经费125 万元
6.	国家自然科学基金 面上项目：喀纳斯湖记录的两万年以来泰加林南缘植被动态与气候变化（课题编号：41571182），2016.1-2019.12，主持，75万

发表文章（*通讯作者）
2023年
1.	Xiang, L., Huang, X.*, Sun, M., Panizzo, V.N.*, Huang, C., Zheng, M., Chen, X. Chen, F. 2023. Prehistoric population expansion in Central Asia promoted by the Altai Holocene Climatic Optimum. Nature Communications. 14, 3102.（SCI-1,Top journal, NI, Nature子刊）
2.	Huang, X.*, Huang, C., Xu, Y., Zheng, M., Ren, X., Chen, X., Hu, Y., Wang, T., Xiang, L., Zhang, J., Chen, F. 2023. Body size of fossil Pediastrum in lake sediments as an indicator of temperature variation. Palaeogeography, Palaeoclimatology, Palaeoecology. 625, 111687.（SCI-2,Top journal）
3.	Cao H., Huang X.*, Xiang, L., 2023. Soil erosion caused the increasing Holocene radiocarbon reservoir effect of Lake Kanas in the Altai Mountains. Radiocarbon. 65, 343-356.（SCI-3, SSCI-1）
4.	Zhang, J., Liu, S., Qiang, M., Chen, Y., Zhang, S., Cheng, B., Pan, X., Li, Y., Huang, X.* 2023. Moisture changes in a mountain–basin system since the last deglaciation revealed by the pollen record of Genggahai Lake, northeastern Qinghai-Tibetan Plateau. Catena. 225, 107046.（SCI-1,Top journal）
5.	Hu, Y., Huang, X.*, Demberel, O., Zhang, J., Xiang, L., Gundegmaa, V., Huang, C., Zheng, M., Zhang, J., Qiang, M., Xiao, J., and Chen F. 2023. Quantitative Reconstruction of Precipitation Changes in the Mongolian Altai Mountains since 13.7 ka. Catena. 107536（SCI-1,Top journal）
6.	Wang, T., Huang, X.*, Zhang, J.*, Luo, D., Zheng, M., Xiang, L., Sun, M., Ren, X., Sun, Y., and Zhang, S. 2023. Vegetation cover dynamics on the northeastern Qinghai-Tibet Plateau since late Marine Isotope Stage 3. Quaternary Science Reviews, 318, 108292.（SCI-1,Top journal）
7.	Ayyamperumal, R.*, Kumari, K., Gandhi, M.S., Huang, X., Chengjun, Z., Nazir, N., Li, F., Das, P. 2023. Environmental hazard assessment and metal contamination in coastal sediments. Chemosphere, 338, 139434. （SCI-2）
8.	Chellaiah, G., Ayyamperumal, R.*, Rengaraj, B., Gopalakrishnan, G., Senapathi, V., Chengjun, Z., Huang, X. 2023. Combined tactic of seasonal changes and ionic processes of groundwater in Tamirabarani river basin, India. Environment Science and Pollution Research, https://doi.org/10.1007/s11356-023-26449-9 （SCI-3）
9.	Li, F.*, Gaillard, M.-J., Cao, X., Herzschuh, U., Sugita, S., Ni, J., Zhao, Y., An, C., Huang, X., Li, Y., Liu, H., Sun, A., Yao, Y. 2023. Gridded pollen-based Holocene regional plant cover in temperate and northern subtropical China suitable for climate modelling. Earth System Science Data, 15, 95-112. （SCI-1,Top journal）
10.	Zhang, D.*, Blyakharchuk, T.A., Bezrukova, E.V., Huang, X., An, C., Su, C., Li, Y. 2023. Holocene rates of vegetation composition change differ between high and middle-to-low elevations in the Central Arid Zone of Asia. Palaeogeography, Palaeoclimatology, Palaeoecology, 618, 111526. （SCI-2,Top journal）

2022年
11.	Ren, X., Huang, X.*, Huang, C., Wang, T., Shen, Z., Zhang, X., Ding, G., Ayyamperumal, R., Zhang, J., Chen, X., 2022. Effects of human activities on mountain forest in northern China during the middle Holocene. Quaternary Science Reviews. 288, 107580.（SCI-1）
12.	Chen, X.*, Huang, X.*, Wu, D.*, Chen, J., Zhang, J., Zhou, A., Dodson J., Atun, Z., Geraldine, J., Yu, J., Wu, Q., Chen, F., 2022. Late Holocene land use evolution and vegetation response to climate change in the watershed of Xingyun Lake, SW China. Catena. 211(3):105973. https://doi.org/10.1016/j.catena.2021.105973. (SCI-1, Top journal)
13.	Li, Y., Huang, X.*, Xiang, L., Zhang, J., Xu, Y., Tian, L., Jin, Y., Jin, X., Zhao, Y., Ren, N., Zhao, H.* 2022. Holocene hydroclimatic change in the Altai Mountains and its impact on human migration. Catena. 219, 106617, https://doi.org/10.1016/j.catena.2022.106617. (SCI-1, Top journal)
14.	Zhang, J., Huang X.*, Qiang, M., Demberel, O., Wang, W., Zheng, M., Xiang, L., Hu, Y., Xiao, J., 2022. Increasing spring insolation in the late Holocene intensified aeolian activity in dryland Asia. Geophysical Research Letters, 17589412, DOI: 10.1029/2022GL101777. (Top journal, Nature Index, SCI-1) 
15.	Zhang, J., Huang X.-Z.*, Wang, J.L., Bradshaw, R.H.W., Wang, T., Xiang, L.X., Luo, D.R., Chen, F.H.*, 2022. An inverse relationship between moisture and grazing intensity in an arid mountain-basin system. Progress in Physical Geography: Earth and Environment. 46(2): 310-322. https://doi.org/10.1177/03091333211060000. (SCI-2).
16.	Li, Y.*, Huang, X.*, Xiang, L., Zhang, J., Xu, Y., Jeppesen, E., 2022. Influence of Lateglacial meltwater influx from alpine glaciers on the aquatic ecosystem of Kanas Lake in the Altai Mountains, China. Palaeogeography, Palaeoclimatology, Palaeoecology. 604(15):  111214. https://doi.org/10.1016/j.palaeo.2022.111214 (SCI-2, Top journal)
17.	Zhang, D., Huang, X., Liu, Q., Chen, X., Feng Z. 2022. Holocene fire records and their drivers in the westerlies-dominated Central Asia. Science of The Total Environment. 833(5):155153 DOI: 10.1016/j.scitotenv.2022.155153. (SCI-1, Top journal)
18.	Zhang, N., Cao, X.*, Xu, Q., Huang, X., Herzschuh, U., Shen, Z., Peng, W., Liu, S., Wu, D., Wang, J., Xia, H., Zhang, D.*, Chen, F. 2022. Vegetation change and human-environment interactions in the Qinghai Lake Basin, northeastern Tibetan Plateau, since the last deglaciation. Catena. 210:105892 (SCI-1, Top journal)
19.	Balasubramanian, M., Sridhar, S.G.D., Ayyamperumal, R.*, Karuppannan, S., Gopalakrishnan, G., Chakraborty, M., Huang, X. 2022. Isotopic signatures, hydrochemical and multivariate statistical analysis of seawater intrusion in the coastal aquifers of Chennai and Tiruvallur District, Tamil Nadu, India. Marine Pollution Bulletin. 174: 113232. (SCI-2).
20.	Dong, G-H, Lu, Y, Zhang, S., Huang, X-Z., Ma, M., 2022. Spatiotemporal variation in human settlements and their interaction with living environments in Neolithic and Bronze Age China. Progress in Physical Geography: Earth and Environment. DOI: 10.1177/03091333221087992 (SCI-2).
21.	Zhao, Y., Miao, Y.*, Huang, X., Chen, C., Zhao, J., Li, Y., Fang, Y., Wang, X., An, C. 2022. Holocene hydroclimate changes revealed by multiple proxies from an alpine lake in the central Tianshan Mountains, Northwest China. Palaeogeography Palaeoclimatology Palaeoecology. 10.1016/j.palaeo.2022.111004. (SCI-2; Top journal)
22.	Cao, X.*, Chen, J., Tian, F., Xu, Q., Herzschuh, U., Telford, R., Huang, X., Zheng, Z., Shen, C. 2022. Long-distance modern analogues bias results in pollen-based precipitation reconstructions. Science Bulletin. DOI: 10.1016/j.scib.2022.01.003 (SCI-1; Top journal)
23.	Fontana, L., Ferreira, P.A., Benassi, R.F., Baldovi, A.A., Figueira, R.C.L., Tambosi, L.R., Calaboni, A., Tavares, D.A., Huang, X., Benassi, S.F., de Souza, J.E., de Jesus, T.A., 2022. Sedimentation rate inferred from 210Pb and 137Cs dating of three sediment cores at Itaipu reservoir (Paraná State, Brazil) the world’s second largest hydroelectricity producer. Journal of Radioanalytical and Nuclear Chemistry. DOI: 10.1007/s10967-022-08380-4 (SCI)
24.	Ayyamperumal, R., Huang, X., Li, F., Zhang, C., Chellaiah, G., Gopalakrishnan, G., Senapathi, V., Perumal, R., Antony, J.K., 2022. Investigation of microplastic contamination in the sediments of Noyyal River- Southern India, Journal of Hazardous Materials Advances, 8, 100198 

2021年
25.	Huang, X.*, Zhang, J., Ren, L., Zhang, S., Chen, F. 2021. Intensification and driving forces of pastoralism in Northern China 5.7 ka ago. Geophysical Research Letters. 48(7), GRL62146. https://doi.org/10.1029/2020GL092288 (SCI-1; Top journal)
26.	Huang, X.*, Xiang, L., Lei, G.*, Sun, M., Qiu, M., Storozum, M., Huang, C., Munkhbayar, C., Demberel, O., Zhang, J., Zhang, J., Chen, X., Chen, J., Chen, F. 2021. Sedimentary Pediastrum record of middle–late Holocene temperature change and its impacts on early human culture in the desert-oasis area of northwestern China. Quaternary Science Reviews. 265, 107054. https://doi.org/10.1016/j.quascirev.2021.107054 (SCI-1; Top journal). 
27.	Huang, X. *, Ren, X., Chen, X., Zhang, J., Zhang, X., Shen, Z., Hu, Y., Chen, F. 2021. Anthropogenic mountain forest degradation and soil erosion recorded in the sediments of Mayinghai Lake in northern China. Catena. 207, 105597. https://doi.org/10.1016/j.catena.2021.105597 (SCI-1; Top journal).
28.	Zhang, J., Huang, X.*, Zhang, S., Ren, X., Xiao, Y., Wang, J., Xiang, L., Xu, Q., Birks, H.J.B. 2021. Cycles of grazing and agricultural activity during the historical period and its relationship with climatic and societal changes in northern China. Land Degradation & Development. 32(11): 3315-3325. https://doi.org/10.1002/ldr.4007 (SCI-1; Top journal, 封面文章)
29.	Peng, W., Huang, X.*, Storozum, M.J., Fan, Y., Zhang, H*. 2021. An updated chronology and paleoenvironmental background for the Paleolithic Loufangzi site, North China. Journal of Human Evolution. 152, 102948. https://doi.org/10.1016/j.jhevol.2020.102948. (Top journal, SCI-2, SSCI-1)
30.	Xiang, L., Chen, X., Huang, C., Sun, M., Xiao, Y., Hu, Y., Huang, X.*, 2021. Divergent patterns of Holocene hydro-climatic evolution in arid central Asia and the Asian summer monsoon margin indicated by Pediastrum records. Palaeogeography, Palaeoclimatology, Palaeoecology. 582, 110662. DOI: 10.1016/j.palaeo.2021.110662. (SCI-2 Top journal)
31.	Xiang, L., Huang, X.*, Huang, C., Chen, X.*, Wang H., Chen J., Hu Y., Sun M., Xiao, Y. 2021. Pediastrum (Chlorophyceae) assemblages in surface lake sediments in China and western Mongolia and their environmental significance, Review of Palaeobotany and Palynology. 289, 104396. https://doi.org/10.1016/j.revpalbo.2021.104396.
32.	Xie, H., Liang, J., Vachula, R.S., Russell, J.M., Chen, C., Guo, M., Wang, X., Huang, X.*, Chen, F. 2021. Changes in the hydrodynamic intensity of Bosten Lake and its impact on early human settlement in the northeastern Tarim Basin, Arid Central Asia. Palaeogeography, Palaeoclimatology, Palaeoecology. 576, 110499. DOI: 10.1016/j.palaeo.2021.110499. (SCI-2, Top journal)
33.	Xiao, Y.1, Xiang, L.1, Huang, X.*, Mills K., Zhang, J., Chen, X., Li, Y. 2021. Moisture changes in the northern Xinjiang basin over the past 2,400 years as documented in pollen records of Jili Lake. Frontiers in Earth Science. 9, 741992. DOI: 10.3389/feart.2021.741992. (1 equal contribution)
34.	Lv, F., Chen, J.*, Zhou, A., Cao, X., Zhang, X., Wang, Z., Wu, D., Chen, X., Yan, J., Wang, H., Dong, G., Xu, Q., Huang, X., Chen, F. 2021. Vegetation history and precipitation changes in the NE Qinghai‐Tibet Plateau: a 7,900-yr pollen record from Caodalian Lake. Paleoceanography and Paleoclimatology. 36(4), e2020PA004126. https://doi.org/10.1029/2020PA004126 (SCI-2, Top journal)
35.	Li, Y., Qiang, M.*, Huang, X., Zhao, Y., Leppänen, J., Weckström, J., Väliranta, M. 2021, Lateglacial and Holocene climate change in the NE Tibetan Plateau: Reconciling divergent proxies of Asian summer monsoon variability. Catena. 199, 105089. https://doi.org/10.1016/j.catena.2020.105089. (SCI-1, Top journal)
36.	Li, Y. *, Hu, L., Wang, H., Zhao, Y., Huang, X., Chen, G., Leppänen, J. J., Fontana, L., Ren, L., Shi, Z., Liu, B., Zhao, H.*, 2021. Meltwater-driven water-level fluctuations of Bosten Lake in arid China over the past 2,000 years. Geophysical Research Letters. 48, 2020GL090988. https://doi.org/10.1029/2020GL090988. (SCI-1, Top journal)
37.	Wang, X., Sun, J., Longstaffe, F.J., Gu, X., Du, S.*, Cui, L., Huang, X., Ding, Z. 2021. Climatic quantification and seasonality of the late MIS 3 in North China: A perspective from carbon and oxygen isotopes of fossil mammal teeth. Quaternary Science Reviews. 272(15): 107222. DOI: 10.1016/j.quascirev.2021.107222. (SCI-1, Top journal)
38.	Ayyamperumal, R.*, Karuppasamy, M.B., Gopalakrishnan, G., Huang, X. 2021. Characteristics of Atmospheric Total Gaseous Mercury Concentrations (TGM) and Meteorological Parameters Observed in Chennai Metropolis, South India. Arabian Journal of Geosciences, 14(1424): 1-11. https://doi.org/10.1007/s12517-021-07803-y.

2020年
39.	Huang, X.*, Sun, M., Xiang, L., Zhang, E., Zhang, J., Grimm, E. 2020. The effect of diatoms on the grain size of lake sediments: a case study of the sediments of Lake Kanas. Journal of Paleolimnology, 63, 101-111. https://doi.org/10.1007/s10933-019-00104-1. (SCI-3)
40.	Huang, X.1*, Zhang J.1, Storozum M., Liu, S., Gill, J.L., Xiang, L., Ren, X., Wang, J., Qiang, M., Chen, F., Grimm, E.C. 2020. Long-term herbivore population dynamics in the northeastern Qinghai-Tibet Plateau and its implications for early human impacts. Review of Palaeobotany and Palynology. 275. 104171. (1 equal contribution; ESI 3%)
41.	Chen, X.*, Wu, D., Huang, X.*, Lv, F., Brenner, M., Chen, F. 2020. Response of vegetation in subtropical southwest China to rapid climate change during the Younger Dryas. Earth-Science Reviews, 201. https://doi.org/10.1016/j.earscirev.2020.103080. (SCI-1, Top journal)
42.	Wang J., Huang, X.*, Zhang, J., Xiang, L., Xiao, Y., Fontana, L., Ren, X., Wang, Z*. 2020. Pollen record of humidity changes in the arid western Qilian Mountains over the past 300 years and comparison with tree-ring reconstructions. Frontiers in Earth Science. 8, DOI: 10.3389/feart.2020.562426.
43.	Li, F., Gaillard, M.-J., Cao, X., Herzschuh, U., Sugita, S., Tarasov, P.E., Wagner, M., Xu, Q., Ni, J., Wang, W., Zhao, Y., An, C., Beusen, A.H.W., Chen, F., Feng, Z., Goldewijk, C.G.M.K., Huang, X., Li, Y., Li, Y., Liu, H., Sun, A., Yao, Y., Zheng, Z., Jia, X., 2020. Towards quantification of Holocene anthropogenic land-cover change in temperate China: A review in the light of pollen-based REVEALS reconstructions of regional plant cover. Earth-Science Reviews. 203, 103119. Doi: 10.1016/j.earscirev.2020.103119. (SCI-1, Top journal)

2019年
44.	Peng, W., Huang, X.*, Zhang, D.J., Storozum, M.J., Chen, F.H., 2019. In-site pollen record from Dadiwan archaeological site and human-environment relationship during Marine Isotope Stage 3. Quaternary Research. 91, 289-300. doi:10.1017/qua.2018.72
45.	Fontana, L., Sun, M., Huang, X. *, Xiang, L., 2019. The impact of climate change and human activity on the ecological status of Bosten Lake, NW China, revealed by a diatom record for the last 2000 years. The Holocene. 2019, 29(12): 1871–1884 DOI: 10.1177/0959683619865586.
46.	Chen, F., Chen, J., Huang, W., Chen, S., Huang, X., Jin, L., Jia, J., Zhang, X., An, C., Zhang, J., Zhao, Y., Yu Z., Zhang, R., Liu, J., Zhou, A., Feng, S., 2019. Westerlies Asia and monsoonal Asia: Spatiotemporal differences in climate change and possible mechanisms on decadal to sub-orbital timescales. Earth-Science Reviews. 192, 337-354.  https://doi.org/10.1016/j.earscirev.2019.03.005. (Top journal, SCI-1, ESI 1%)
47.	黄小忠，向丽雄，张恩源，孙明杰. 2019. 全新世中期7 ka前后降温事件对中国北方植被生态的影响. 第四纪研究. 39（3）：687-700. Doi: 10.11928/j.issn.1001.7410.2019.03.
48.	周刚平，黄小忠*，王宗礼，张军，谢海超. 2019. 基于粒度数据重建的近2000a新疆博斯腾湖区域风沙活动. 中国沙漠，39（2）：86-95.
49.	黄小忠，勾晓华. 2019. 高校“中国与世界地理” 公选课的课程架构及教学探讨. 高等理科教育，145：108-113.  【教学论文】

2018年
50.	Huang X.Z.*, Peng W., Rudaya N., Grimm E.C., Chen X.M., Cao X.Y., Zhang J., Pan X.D., Liu S.S., Chen C.Z., Chen F.H. 2018. Holocene vegetation and climate dynamics in the Altai Mountains and surrounding areas. Geophysical Research Letters. 45, 6628-6636. DOI: 10.1029/2018GL078028. (SCI-1，Top journal; ESI 5%)
51.	Huang, X.*, Chen, X., Du, X., 2018. Modern pollen assemblages from human-influenced vegetation in northwestern China and their relationship with vegetation and climate. Vegetation History and Archaeobotany. 27(6): 767-780. DOI: 10.1007/s00334-018-0672-0.
52.	Zhang, J., Huang, X.*, Wang, Z., Yan, T., Zhang, E. 2018. A late-Holocene pollen record from the western Qilian Mountains and its implications for climate change and human activity along the Silk Road, Northwestern China. The Holocene. 28(7), 1141-1150. DOI: 10.1177/0959683618761548.
53.	Lin, X., Rioual, P.*, Peng, W., Yang, H., Huang, X.* 2018. Impact of recent climate change on Lake Kanas, Altai Mountains (N.W. China) inferred from diatom and geochemical evidence. Journal of Paleolimnology. 59(4), 461-477. https://doi.org/10.1007/s10933-018-0019-y.
54.	Zhang, J., Xia, Z.*, Zhang, X., Storozum, M.J., Huang, X.*, Han, J., Xu, H., Zhao, H., Cui, Y., Dodson, J., Dong, G. 2018. Early-middle Holocene ecological change and its influence on human subsistence strategies in the Luoyang Basin, north-central China. Quaternary Research. 89(2), 446-458. DOI:10.1017/qua.2017.104.
55.	Xie, H., Zhang, H., Ma, J., Li, G., Wang Q, Rao, Z., Huang, W., Huang, X., Chen, F. 2018. Trends of increasing Holocene summer precipitation in arid central Asia: evidence from an organic carbon isotopic record from LJW10 loess section in Xinjiang, NW China. Palaeogeography, Palaeoclimatology, Palaeoecology, 509, 24-32. DOI: 10.1016/j.palaeo.2018.04.006.
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